The prosperous land of Kumandra is ravaged by the Druun, evil spirits that attack and petrify the land's people and dragons. Sisu, the last dragon, concentrates her magic into a gem and banishes the Druun, reviving Kumandra's people but not the dragons. A power struggle for the orb divides Kumandra's people into tribes, based on their placement along a giant, dragon-shaped river: Fang, Heart, Spine, Talon, and Tail.
500 years later, Chief Benja of the Heart tribe trains his daughter, Raya, until she finally becomes a guardian of the gem. Firmly believing that the tribes can be united once again, Chief Benja decides to hold a feast for the leadership of all five tribes. During the feast, Raya befriends the daughter of Chief Virana of the Fang tribe, Namaari, who gives Raya a dragon pendant as a gift. Trusting her, Raya shows Namaari the location of the gem but Namaari, betraying Raya, reveals that she is trying to help Fang steal it. Alerted to the attack, Benja and the other tribes arrive at the scene and start fighting over the gem, which gets destroyed in the scuffle. As each tribe steals a piece of the orb, the reawakened Druun ravage Heart and its people. A wounded Chief Benja, discovering that the Druun are weak against water, throws Raya into a river before getting petrified himself.
For the next six years, Raya treks across Kumandra to find Sisu in order to help her recover the missing orb pieces. After reaching a ship wreck in Tail, Raya finally manages to summon Sisu, who admits that she did not create the orb, but only used it on behalf of her siblings and can gain her siblings' powers when she touches its pieces.
They recover the second piece from a ruined, booby-trapped Tail temple, granting Sisu shapeshifting powers; and encounter a hostile Namaari and her tribesmen after Raya's stolen portrait of Sisu. Escaping the temple, they escape on the boat of young restauranteur Boun, who lost his family to the Druun. Despite Boun's friendliness, Raya does not fully trust him, and insists that Sisu stays in human form.
The group arrives at Talon to claim the third gem piece, during which Raya encounters "con baby" Noi and her trio of monkey-like companions, the Ongis, who adopted her after the Druun petrified her mother. After a chase, Raya recruits Noi and the Ongis while Sisu encounters Talon's chieftess, who sinisterly offers her to the Druun for the other orb pieces. Raya rescues Sisu and reclaims the third piece, allowing Sisu to spew out fog which they use to escape.
The group then arrives at Spine, where they meet Tong, a fearsome warrior, and the village's sole survivor. Namaari arrives and Raya holds her off in a fight so the others can escape, but before Namaari is able to defeat her, Sisu turns back into a dragon and saves Raya. Realizing that Sisu is a dragon, the companions agree to help Raya, and Tong hands over Spine’s gem piece, allowing Sisu to control rain.
As the group get closer to Fang, Sisu suggests allying with Namaari rather than stealing the final piece. When Raya refuses, Sisu takes her back to the remains of the Heart and tells of how her siblings' (Pangu, Jagan, Pranee, and Amba) trust in her was what really empowered her to save Kumandra. Raya relents and decides to give Namaari the dragon pendant as a peace offering. Raya and Sisu meet privately with Namaari, but Namaari, torn by her responsibility to save Fang's reputation and her wish to help defeat the Druun, threatens to shoot them with a crossbow upon seeing the orb pieces. Sisu tries to calm Namaari down but is fatally shot when Raya lashes her sword on Namaari's crossbow.
With the last dragon dead, and the water subsequently gone, Fang is overrun by the Druun. Raya enters to confront Namaari, whom she finds to be mourning over the petrification of her mother. The two fight while Raya's companions rescue the people of Fang using the gem pieces. Raya, besting Namaari, prepares to kill her, but stops upon realizing her fault in Sisu’s death due to her inability to trust others and heads off with Namaari to aid the group instead. As the Druun gain on Raya's group, Raya, remembering how trust allowed Sisu to save the world beforehand, urges the others to unite and reassemble the gem, showing her faith towards Namaari by handing her piece and allowing the Druun to take her. The rest follow suit, with Namaari assembling the gem before the Druun petrify her as well.
With the gem reassembled, and everybody trusting each other, the Druun are vanquished and everyone, including the dragons, are revived. The dragons revive Sisu while everyone reunites with their lost loved ones, including Raya and Chief Benja. The dragons and the tribes gather at Heart to celebrate and finally reunite as Kumandra.
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